N A T U R K U N D L I C H E S   I N F O R M A T I O N S S Y S T E M

Micropterix tuscaniensis Heath, 1960

(zoological nomenclature: valid name, available)

General information:

Micropteryx tuscaniensis Heath, 1960: 130 - 132.
Type locality: Italy, Tuscany, Pratolino
Type: Holotype : Pratolino, Tuscany, 29.IV.1893 Wlsm. 85691 (genitalia investigation 7083 J.F.G.C.); in coll. BMNH, London

Synonyms, misspellings, wrong determinations, etc.:
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Habit:

   
: Tuscany, Alpi Apuane, near Massa, Campareccia, 1992.05.30, leg. Hausenblas/Zeller/Kurz, coll. Michael Kurz : Tuscany, Alpi Apuane, near Massa, Campareccia, 1992.05.30, leg. Hausenblas/Zeller/Kurz, coll. Michael Kurz    
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
   

Scheme of wing pattern:

Description of adults: Examined: 9 , 6 . Forewing length: 3.4 - 4.3 mm; 3.7 - 4.7 mm. Head black-brown; vestiture of hair-like scales on the head dirty to rusty yellow; antennae dark brown, bronzy golden shining, sometimes slightly tinged purple, 3/4 (), respectively, slightly more than 1/2 () of forewing length; thorax bronzy golden, distally somewhat purple; tegulae purple to purple-violet, distally sometimes with blueish scales; forewings purple to purple-violet, medially also tinged blueish; basis of wing sometimes slightly bronzy golden; markings whitish golden to golden, delicately bordered bronzy: At nearly 1/4 a somewhat variable fascia, nearly tapering to one point at costa, very broad at inner margin and therefore more or less triangle-shaped; an irregular, outwardly bent fascia across the whole wing width at almost 1/2; a small costal spot at 3/5; a variable spot at 3/4, mostly reaching from costa across half of the wing width; tip of wing and outer margin often coppery; fringe bronzy golden, at its basis sometimes purplish tinged, whitish outwards; hindwings bronzy golden, more or less tinged purple, especially apically; fringe bronzy golden with purplish tinge, whitish outwards; legs and abdomen brown, golden shining.

Remarks: The determination of the two smallest females, on which the above description is partly based, is uncertain.

Distribution:

Following Heath (1960b), the species has been found only in Italy, in the provinces of Tuscany, Calabria, Puglia and Basilicata. The investigated specimens are from the vicinity of Triest, Emilia, Tuscany and Lazio, but the determination of the two specimens from Triest and Lazio is uncertain. Most probably, M. tuscaniensis is distributed throughout the whole of mainland Italy, except the Alps. The vertical distribution reaches from near sea level to about 1050 m a.s.l. (Kurz & Kurz 2010).

Biology:

In Tuscany, the animals have been found frequently at the edge of dense shrub- and woodland at noon. The biotop was situated at a rocky slope in full sunshine. There, the animals sat underneath the leaves of beech, where they were crawling and probably feeding on pollen, gathered on the surface of the young, somewhat sticky leaves. Furthermore, they have also been found on blossoms of Sambucus nigra. The imagines are on the wing from late April to early July, depending on the altitude (Kurz & Kurz 2010).

Stages in development:

The early stages are unknown.

Anatomy:

, scheme of cuticle -genitalia, prep.-no. 894, M. Kurz: Tuscany, Alpi Apuane, near Massa, Campareccia, 1992.05.30, leg. H/Z/K, coll. Michael Kurz -genitalia, prep.-no. 945, M. Kurz: Tuscany, Firenze, Marradi, Ponte Valle, 500 m, 2003.06.20, coll. Museo Friulano di Storia Naturale, Udine -genitalia, prep.-no. 525, M. Kurz: "EMILIA; Saibola [?]; 5.6.[18]92; A. FIORI", coll. Naturhistorisches Museum, Vienna
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
-genitalia, aedeagus in higher magnification, prep.-no. 525, M. Kurz: "EMILIA; Saibola [?]; 5.6.[18]92; A. FIORI", coll. Naturhistorisches Museum, Vienna , scheme of cuticle -genitalia, preparation in caustic potash, prep.-no. 895, M. Kurz: Tuscany, Alpi Apuane, near Massa, Campareccia, 1992.05.30, leg. H/Z/K, coll. Michael Kurz -genitalia, prep.-no. 895, M. Kurz: Tuscany, Alpi Apuane, near Massa, Campareccia, 1992.05.30, leg. H/Z/K, coll. Michael Kurz
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
   
-genitalia, prep.-no. 330, M. Kurz [identity uncertain]: "Italia, Terrelle, M[onte]; Cassino, 24.6.1970; 900 m J.Klimesch", coll. Zoologische Sammlung des Bayrischen Staates, Munich -genitalia, receptaculum seminis in higher magnification, prep.-no. 895, M. Kurz: Tuscany, Alpi Apuane, near Massa, Campareccia, 1992.05.30, leg. Hausenblas, Zeller, Kurz, coll. Michael Kurz    
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
   

genitalia. Uncus moderately long, very broad and stout, with broadly rounded tip; ventrally of the uncus, inside the tegumen a structure with long, hair-like setae; between uncus and accessory claspers, at the distal margin of the tegumen, two rows of short and thin, nevertheless stout and somewhat thickened setae with spatulate tips; accessory claspers broad, putty-like, at the margin opposite to the valves at the inner surface a row of about 6 long, distally partly twisted, stout and thickened setae; proximad of this row a small appendix with a single long, distinctly twisted, stout and thickened seta; above the first row a second one of about 5 modified, T-shaped, and 2 - 3 smaller, bent, but otherwise unmodified, stout and thickened setae; valves moderately long, somewhat stout, constricted distad of the middle, the distalmost fourth strongly spoon-like inflated; at the inner surface of the valves, postbasally a row of short setae, at about 1/3 of their length a single, stout and thickened seta; the distal fourth of the valves at the inner surface bearing numerous long, stout and thickened setae.

genitalia. Tergite IX missing; sternite IX strongly reduced, weakly sclerotized, with indistinct lateral margins; papillae anales with band-like sclerotization, but of no diagnostic value; receptaculum seminis short, the proximal 2/3 bag-like inflated, distinctly striated.

Diagnosis:

M. tuscaniensis : Tuscany, Alpi Apuane, near Massa, Campareccia, 1992.05.30, leg. Hausenblas/Zeller/Kurz, coll. Michael Kurz M. cassinella ALLOTYPE: IT, Monte Cassino, Terelle, 1970.06.24, leg. Josef Klimesch; coll. Zoologische Sammlung des Bayrischen Staates, Munich (ZSM) M. hartigi : Basilicata, Lucano, Viggianello, 1000 m, 2000.06.07, leg. Zeller, coll. Michael Kurz M. allionella : Piemonte, Cuneo, Valle Corsaglia, 1050 m, 1973.05.25, leg. Jaeckh, coll. United States National Museum, Washington
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
M. zangheriella : Emilia Romagna, near Lizzano in Belvedere, 760 m, 2002.05.30, leg. Renate Hausenblas und Christof Zeller, coll. Christof Zeller M. tuscaniensis -genitalia, prep.-no. 894, M. Kurz: Tuscany, Alpi Apuane, near Massa, Campareccia, 1992.05.30, leg. H/Z/K, coll. Michael Kurz M. cassinella -genitalia, prep.-no. 765 M. Kurz: PARATYPE; Lazio, Frosinone, Capo d´Acqua, 1200 m, 1982.07.08, leg. Josef Klimesch, coll. Michael Kurz M. hartigi, -genitalia, prep-no. 676, M. Kurz: Puglia, Promontorio del Gargano, Foresta Umbra, 1993.05.22, leg. Zeller/Kurz, coll. Michael Kurz
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
M. allionella -genitalia, prep.-no. 750, M. Kurz: Liguria, Savona, Noli, 1951.04.26, leg. Josef Klimesch, coll. Michael Kurz M. zangheriella -genitalia, prep.-no. 944, M. Kurz: Toscana, Firenze, Marradi, Badia Valle, 430 m, 1997.05.13, coll. Museo Friulano M. tuscaniensis -genitalia, prep.-no. 895, M. Kurz: Tuscany, Alpi Apuane, near Massa, Campareccia, 1992.05.30, leg. H/Z/K, coll. Michael Kurz M. cassinella -genitalia, prep.-no. 759 M.Kurz: Campania, San Salvatore Telesino, 2000.06.10, leg. Lastuvka, coll. Jan Liska
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
 
M. hartigi -genitalia, prep.-no. 887, M. Kurz: Basilicata, Lucano, Viggianello, 1000 m, 2000.06.07, leg. Christof Zeller, coll. Michael Kurz M. allionella -genitalia, prep.-no. 812, M. Kurz: Emilia Romagna, northern Apennines, Sestola, 1992.05.29, leg. Hausenblas, Zeller, Kurz, coll. Michael Kurz M. zangheriella -genitalia, prep.-no. 954, M. Kurz: Emilia Romagna, near Lizzano in Belvedere, 760 m, 2002.05.30, leg. C. Zeller, coll. Michael Kurz  
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
 

Externally, M. tuscaniensis can be distinguished from other sympatric species without golden markings on inner and outer margin best by the form of the fascia at 1/4, which is broad triangle-shaped. Micropterix cassinella Kurz, Kurz & Zeller, 2010 usually is smaller and, like Micropterix hartigi Heath, 1981, often has a less violet ground colour. Nevertheless, some females of M. tuscaniensis and M. cassinella may be indistinguishable. In contrast to M. hartigi, but also to Micropterix allionella (Fabricius, 1794), the spot at 3/4 is distinctly smaller in M. tuscaniensis.
In the male genitalia, there aren´t any similar species to M. tuscaniensis, in the female genitalia on the other hand, no convincing diagnostic features have been found so far.

Phylogeny: The systematic position of M. tuscaniensis within the genus is still unclear. Morphologically, the male genitalia show some similarities to Micropterix cassinella Kurz, Kurz & Zeller, 2010 on the one hand (enlargement of the distal portion of the valves), and to Micropterix zangheriella Heath, 1963 and Micropterix hartigi Heath, 1981 on the other hand. In the latter case, all three species have the basalmost thickened seta on the ventral margin of the accessory claspers distinctly offset from the rest of the row and strongly modified. in M. cassinella too, this seta is offset, but the accessory claspers are generally somewhat simpler, with less modified setae. Wether these similarities indicate a closer relationship however, remains uncertain due to the lack of additional, especially DNA-data.

Worth knowing:

Sources:

Heath, J. 1960b. Two new species of Micropteryx (Lepidoptera: Micropterygidae); Estratto dal Bolletino della Società Italiana, Vol. XC, no. 7 - 8: 129 - 133.
Kurz, M. A. & M. E. Kurz 2000–2010. Naturkundliches Informationssystem. – URL: http://www.nkis.info [online 05 November 2010].

Publication data:

history:
Kurz Michael: 2009.02.11
Kurz Michael: 2010.01.25
Kurz Michael: 2010.02.16
Kurz Michael: 2010.10.12
Kurz Michael: 2010.12.12
Kurz Michael: 2010.12.15
Kurz Michael: 2010.12.16
Kurz Michael: 2011.12.17
Kurz Michael: 2012.02.12
Kurz Michael: 2012.03.23
Document reviewed by:
not reviewed: 2012.03.23
Document released by:
Kurz Michael: 2012.03.23

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