N A T U R K U N D L I C H E S   I N F O R M A T I O N S S Y S T E M

Alloclemensia mesospilella (Herrich-Schäffer, [1853])

(zoological nomenclature: valid name, available)

General information:

Incurvaria mesospilella Herrich-Schäffer, [1853]: 67.
Type locality: Austria, Lower Austria, Semmering and Rax
Synonyms, misspellings, wrong determinations, etc.:
Incurvaria trimaculella Herrich-Schäffer 1854: Junior homonym of Incurvaria trimaculella Duponchel, 1840 (now Telechrysis tripuncta (Haworth, 1828), Amphisbatidae)
Lampronia Trimaculella v. Quadrimaculella Höfner 1900: 251. Type locality: Carinthia, Koralpe. Junios subjective synonym of Incurvaria mesospilella.

Habit:

   
: Trentino, Monte Baldo, Bocca di Navene, 1500 m, 1970.07.15, leg. K. Burmann, coll. Tiroler Landesmuseum Ferdinandeum : Trentino, Monte Baldo, Bocca di Navene, 1500 m, 1970.07.15, leg. K. Burmann, coll. Tiroler Landesmuseum Ferdinandeum    
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
   

Description of adults: Examined: 2 , 1 . Forewing length 5.5 - 5.9 mm, 5.3 mm. Neck and vertex distantly scaled, ochreous; frons more beige, less distantly scaled; diameter of eye about 0.25 mm, distance of eyes (frontal) nearly twice the diameter; maxillary palpi well developed, with adjacent scales, beige to ochreous; labial palpi nearly twice as long as the diameter of an eye, with adjacent scales, anteriorly directed downwards, slightly diverging; antennae filiform, about 1/2 of forewing length in both sexes, bearing beige scales and fine cilia; thorax and tegulae brown, bronzy golden shining; forewings elongate with acute apex, of coffee- to dark brown colour, with a strong bronzy golden glance; at about 1/4 a more or less moderately broad fascia across the whole wing width, whitish with bronzy golden shimmer; basalwards of the tornus a broad, triangular, whitish spot, its tip reaching across 1/3 of the wing width; on the costal margin, distally of 1/2 a whitish, nearly triangular spot reaching nearly across 1/3 of the wing width, somewhat displaced basalwards compared with the tornal spot; sometimes a small whitish, elongate spot at the costal margin at about 4/5; fringe dark brown from costal margin to apex, along the outer margin distinctly bicoloured, whitish distally, brown with bronz ygolden shimmer proximally; at the tornus, fringe brown again, bronzy golden shining; hindwings with broad scales, completely covering the wing membrane, greyish brown with nacre-like shimmer to dark brown, shimmering purple-violet with single blueish scales; fringe brown, bronzy golden shining; underside similar in colouration to the upperside, on the forewings light markings only very diffuse; fringe similar too; legs brown, bronzy golden shining; foretibia with an epiphysis, midtibia with a pair of spurs at the distal end; hindtibia thickened, with a tuft of long, hair-like scales and a pair of spurs each, distalwards of its middle and at the distal end; abdomen greyish brown, bronzy golden shining.

Distribution:

A. mesospilella has been recorded from central Europe (especially from the Alps), from northern Europe, but also from the Balkans (Bosnia, Romania). The species is absent from the British Isles and from most of the Mediterranean region (Karsholt & Nieukerken 2004). It has been recorded from North America too by Hodges et al. 1983, but these records refer to Alloclemensia americana Nielsen, 1981. Vertical distribution has been documented from near sea-level in Denmark to 1800 m a.s.l. in the Alps (Kurz & Kurz 2018).

Biology:

The species inhabits shady woods, ravines, but also elfin woodland. The imagines are on the wing in one generation per year from May to July (Kurz & Kurz 2018).

Stages in development:

 
Early mine on Saxifraga rotundifolia: Salzburg, Osterhorn group, Hintersee, Satzstein, 2018.09.28 Holes from cutting out the leaf bag: Salzburg, Osterhorn group, Hintersee, Satzstein, 2018.09.28 Salzburg, Osterhorn group, Hintersee, Satzstein, 2018.09.28  
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
 

Larva. Following Meeß (in Spuler 1910), the larva is yellowish white with black head and prothoracic plate. It first mines in the leaves of Saxifraga rotundifolia. The mine is at first a slender galery, later it becomes a blotch. The larva cuts out a roundish piece of the leaf and feeds beneath it, constructing a flat case from the remnants. From time to time, it adds further leaf-pieces. In autumn, it drops down to the ground in a now oval case, hides preferably between mosses and pupates.

Pupa. The pupa hibernates in its case.

Anatomy:

-genitalia, prep.-no. 865, M. Kurz: Trentino, Monte Baldo, Bocca di Navene, 1500 m, 1970.07.15, leg. Karl Burmann, coll. Tiroler Landesmuseum Ferdinandeum, Innsbruck -genitalia, prep.-no. 865, M. Kurz, uncus and valve in higher magnification: Trentino, Monte Baldo, Bocca di Navene, 1500 m, 1970.07.15, leg. Karl Burmann, coll. Tiroler Landesmuseum Ferdinandeum, Innsbruck -genitalia, prep.-no. 869, M. Kurz, uncus in higher magnification: Trentino, Monte Baldo, Bocca di Navene, 1500 m, 1970.07.15, leg. Karl Burmann, coll. Tiroler Landesmuseum Ferdinandeum, Innsbruck -genitalia, prep.-no. 869, M. Kurz, valve in higher magnification: Trentino, Monte Baldo, Bocca di Navene, 1500 m, 1970.07.15, leg. Karl Burmann, coll. Tiroler Landesmuseum Ferdinandeum, Innsbruck
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
-genitalia, prep.-no. 865, M. Kurz, aedoeagus in higher magnification: Trentino, Monte Baldo, Bocca di Navene, 1500 m, 1970.07.15, leg. Karl Burmann, coll. Tiroler Landesmuseum Ferdinandeum, Innsbruck -genitalia, prep.-no. 869, M. Kurz, aedoeagus in higher magnification: Trentino, Monte Baldo, Bocca di Navene, 1500 m, 1970.07.15, leg. Karl Burmann, coll. Tiroler Landesmuseum Ferdinandeum, Innsbruck -genitalia, prep.-no. 537, M. Kurz: Salzburg, Radstädter Tauern, Tappenkarsee, 1990.07.15, leg. Heimo Nelwek, coll. Michael Kurz -genitalia, tip of ovipositor, prep.-no. 537, M. Kurz: Salzburg, Radstädter Tauern, Tappenkarsee, 1990.07.15, leg. Heimo Nelwek, coll. Michael Kurz
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
Picture from: Kurz Michael
Detailed view
     
Foretibia with epiphysis, prep.-no. 934, M. Kurz: Trentino, Monte Baldo, Bocca di Navene, 1500 m, 1970.07.15, leg. Karl Burmann, coll. Tiroler Landesmuseum Ferdinandeum, Innsbruck      
Picture from: Kurz Michael
Detailed view
     

genitalia. Posterior part of ovipositor triangular, tip pronounced; lateral margins with two steps, the anterior steps distinct; apophyses posteriores and anteriores both relatively long, posterior ones distinctly longer than anterior ones; ductus bursae long, sclerotized in its first part, with two convolutions; bursa copulatrix bag-like, of moderate size, without signa.

Diagnosis:

In Europe, the species can readily be distinguished from similar species by the usually lighter brown ground colour and strong bronzy golden reflections on the forewing upperside. Specimens with an additional preapical spot may be mistaken for Lampronia rupella ([Denis & Schiffermüller], 1775), although the latter normally has more prominent white spots and a darker ground colour. In the male genitalia, A. mesospilella can readily be recognized by the finger-like process of the uncus and by the shape of the valvae, whereas in the female genitalia, the species lacks the stellate signa in the bursa copulatrix.

Worth knowing:

Sources:

Duponchel, P. A. J. [1838–1840]. Nocturnes, 8. – In: J.-B. Godart, Histoire Naturelle des Lépidoptères ou Papillons de France 11. – Paris, Méquignon-Marvis. 720 pp., pls 287–314.
Haworth, A. H. 1828. Lepidoptera Britannica, sistens digestionem novam insectorum lepidopterorum quae in Magna Britannia reperiuntur, lavarum pabulo, temporeque pascendi; expansione alarum; mensibusque volandi; synonymis atque locis observationibusque variis. Part 4. Londini, pp. 513–609.
Herrich-Schäffer, G. A. W. 1847–1855 ["1853–1855"]. Systematische Bearbeitung der Schmetterlinge von Europa, zugleich als Text, Revision und Supplement zu Jakob Hübner’s Sammlung europäischer Schmetterlinge. 5. Die Schaben und Federmotten: [1]–2–394 + [1]–2–52, 124 + 7 + 1 pl. Regensburg.
Hodges, R. W. et al. 1983. Check List of the Lepidoptera of America North of Mexico. E. W. Classey, London, UK. Updated by Jean-François Landry, Don Lafontaine and Jim Troubridge [available on http://www.lepbarcoding.org/cl_nth_am.php]
Höfner, G. 1900. Die Schmetterlinge des Lavantthales und der beiden Alpen Kor- und Saualpe - Jahrbuch des Naturhistorischen Landesmuseums von Kärnten 26: 247-253.
Karsholt, O. & E. J. van Nieukerken 2004. Incurvariidae. In - Karsholt, O. & E. J. van Nieukerken (eds.). Lepidoptera, Moths. – Fauna Europaea version 1.1, http://www.faunaeur.org [online 3 July 2008].
Kurz, M. A. & M. E. Kurz 2018. Naturkundliches Informationssystem. - URL: http://www.nkis.info [online 2018.10.07].
Meeß, A. 1910. Tineides aculeatae. - in A. Spuler: Die Schmetterlinge Europas, Kleinschmetterlinge; unveränderter Nachdruck der S. 188-523 und der Tafeln 81-91, Verlag E. Bauer, Keltern, 1983.
Nielsen, E. S. 1981. A taxonomic revision of the species of Alloclemensia n.gen. (Lepidoptera: Incurvariidae s. str.). Insect Systematics & Evolution 12 (3): 271–294.

Publication data:

history:
Kurz Michael: 2009.09.08
Kurz Michael: 2011.12.28
Kurz Michael: 2012.11.02
Kurz Michael: 2017.03.31
Kurz Michael: 2017.04.26
Kurz Michael: 2018.09.19
Kurz Michael: 2018.09.20
Kurz Michael: 2018.10.07
Kurz Michael: 2018.10.31
Kurz Michael: 2018.11.15
not reviewed

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